Twitter still uses Ruby on Rails to generate the website, as always. Their use of Scala is on the backend; the services that lay behind the Rails app. Facebook uses a similar architecture behind their PHP code, using many other languages, including Java.
Almost no one is facing the same issues as twitter. I find it extremely unlike that DHH thinks Rails is best for every possible scenario, and a single counterexample of an extreme outlier isn't an argument against this.
[1] twitter dot com