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From an evolutionary standpoint, they might as well never have lived

There is the distinct possibility that they affected the chance for others to reproduce or survive. So you are right about their individual evolution, but they were likely part of group or species evolution.

In bees and other hive insects this effect is most pronounced, as there are only only a few individuals in the hive that can reproduce. But with only them, the hive would die out, so the rest is useful from an evolutionary standpoint.




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