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Two examples of rapid prototyping spring to mind:

* Twitch, started as one guy streaming his life then they realised lots of gamers were watching, and that they'd like to be able to stream https://www.youtube.com/watch?v=FBOLk9s9Ci4

* Segment, started as a thumbs up/down tool for professors in lectures to work out when students are getting confused. They realised everyone just went to Facebook instead, then they wondered why they couldn't tell this when they were remote! https://www.youtube.com/watch?v=l-vfn97QTr0

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