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first they argue that the MOAD and Xerox' Smalltalk pioneered dark-on-light after years of the reverse, then they argue that actually it's because vision-psychology. Well, which is it? Because there's no way the MOAD and Smalltalk chose dark-on-light because of visual perception research. Smalltalk's choice was probably motivated by a desire to make the UI look like paper (Xerox being in the paper business). Same with Apple's original Mac. At the time everyone's UI was paper so this familiar UI was important to replicate to sell.

It also ignores the fact that we've had light-on-dark mode for ages: blueprints and microfiche, blackboards and granite tablets

I mean, the Rosetta Stone is light-on-dark

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